Here is a list of publications from our lab since 2006 (and a few earlier ones).  If the PDF is not posted, please contact me.  We’re trying to publish mainly in journals that operate on a non-profit basis.  Asterisks indicate student authors.


Vickruck JL*, MH Richards.  A test of kin and nestmate recognition in the eastern carpenter bee.  Behavioural Processes 145, 73-80.  Click here for free copy until 6 December!

Onuferko TM*, DA Skandalis*, R Leon Cordero*, and MH Richards. Rapid initial recovery and long-term persistence of a bee community in a former landfill. Insect Conservation and Diversity, Early Online version, doi: 10.1111/icad.12261.

Vickruck JL*, Richards MH. Nesting habits influence population genetic structure of a bee living in anthropogenic disturbance. Molecular Ecology, 26(10), 2674-2686, doi:10.1111/mec.14064

Lewis V*, MH Richards.  Experimentally induced alloparental care in a solitary carpenter bee. Animal Behaviour 123 (2017) 229-238.

Hudson LN et al.  The database of the PREDICTS (Projecting Responses of Ecological Diversity In Changing Terrestrial Systems) project.  Ecology and Evolution 7(1), 145-188.  DOI: 10.1002/ece3.2579


De Palma A. et al. Predicting bee community responses to land-use changes: Effects of geographic and taxonomic biases. Scientific Reports 6, 31153; doi: 10.1038/srep31153


Onuferko TM, R Kutby, and Richards MH.  A list of bee species (Hymenoptera: Apoidea) recorded from three municipalities in the Niagara region of Ontario, including a new record of Lasioglossum furunculum Gibbs (Halictidae) in CanadaJournal of the Entomological Society of Ontario 146: 3–22

Richards MH, C Course. Ergonomic skew and reproductive queuing based on social and seasonal variation in foraging activity of eastern carpenter bees. Canadian Journal of Zoology, 93(8): 615-625, DOI 10.1139/cjz-2014-0330 (Reviews round 1 and round 2)

Nonacs P, MH Richards.  How (not) to review papers on inclusive fitness.  Trends in Ecology and Evolution 30(5), 235-237.  DOI:

Richards MH, TM Onuferko, SM Rehan.  Phenological, but not social, variation associated with climate differences in a eusocial sweat bee, Halictus ligatus, nesting in southern Ontario.  Journal of Hymenoptera Research 43, 19-44.  DOI: 10.3897/JHR.43.8756   PDF


Vickruck JL.  Development of sixteen novel microsatellite markers for the eastern carpenter bee, Xylocopa virginica (Hymenoptera: Apidae), through paired-end Illumina sequencing.  Conservation Genetics Resources (2015) 7:427–429.  DOI 10.1007/s12686-014-0387-1

Rehan SM, MH Richards, MH Adams, MP Schwarz. The costs and benefits of sociality in a facultatively social bee. Animal Behaviour 97, 77-85. DOI


Rutgers-Kelly A, Richards MH. 2013. Effect of meadow regeneration on bee (Hymenoptera: Apoidea) abundance and diversity in southern Ontario, Canada.  The Canadian Entomologist145, 06, 655-667, DOI: 10.4039/tce.2013.42.    PDF (Kudos: Cambridge University Press)

Rehan SM, A Rotella, TM Onuferko, MH Richards. 2013. Colony disturbance and solitary nest initiation by workers in the obligately eusocial sweat bee, Halictus ligatus.  Insectes Sociaux 60, 3, 389-392, DOI: 10.1007/s00040-013-0304-8

Rehan SM, MH Richards. 2013. Reproductive aggression and nestmate recognition in a subsocial bee. Animal Behaviour 85, 733-741, DOI: 10.1016/j.anbehav.2013.01.010


Vickruck JL, MH Richards. 2012. Niche partitioning based on nest site selection in Ceratina mikmaqi and C. calcarata. Animal Behaviour 83, 1083-1089.


Rehan SM, MP Schwarz, MH Richards. 2011. Fitness consequences of ecological constraints and implications for the evolution of sociality in an incipiently social bee. Biological Journal of the Linnean Society 103, 57-67.

Richards MH. 2011. Colony social organisation and alternative social strategies in the eastern carpenter bee, Xylocopa virginica. Journal of Insect Behavior 24, 399-411.

Richards MH, A Rutgers-Kelly, J Gibbs, JL Vickruck, SM Rehan, C Sheffield. 2011. Bee diversity in naturalizing patches of Carolinian grasslands in southern Ontario. The Canadian Entomologist 143: 279–299.  PDF

Vickruck JL, SM Rehan, CS Sheffield, MH Richards. 2011. The nesting biology of two cryptic species of Ceratina dupla, with comparisons to C. calcarata (Hymenoptera: Apidae). The Canadian Entomologist 143, 254–262.  PDF

Skandalis DA, MH Richards, S Todd, G Tattersall. 2011. Cold tolerance and its influence on breeding phenology and geographic range of the large carpenter bee, Xylocopa virginica. Canadian Journal of Zoology 89, 785-795.


Peso M, MH Richards. 2010. Knowing who’s who: nestmate recognition in the facultatively social carpenter bee, Xylocopa virginica. Animal Behaviour 79, 563-570.

Rehan SM, MH Richards, MP Schwarz. 2010. Social polymorphism in the Australian small carpenter bee, Ceratina (Neoceratina) australensis. Insectes Sociaux 57, 403-412.

Rehan SM, MH Richards. 2010. Nesting biology and subsociality in the small carpenter bee, Ceratina calcarata (Hymenoptera: Apidae). The Canadian Entomologist 142, 65-74.  PDF

Sheffield CS, T Griswold, MH Richards. 2010. Discovery of the Old World bee, Megachile (Pseudomegachile) ericetorum (Hymenoptera: Megachilidae), in Ontario, Canada. Journal of the Entomological Society of Ontario 141, 85-92.

Richards MH, L Packer. 2010. Social behaviours in solitary bees: Interactions among individuals in Xeralictus bicuspidariae Snelling (Hymenoptera: Halictidae: Rophitinae). Journal of Hymenoptera Research 19, 66-76.

Vickruck JL, J Huber, MH Richards. 2010. Nest parasitoids of the bee genus Ceratina (Hymenoptera: Apidae) in the Niagara Region, Ontario, Canada. Journal of the Entomological Society of Ontario, 141, 11-26.

Peso M and MH Richards. 2010. Not all who wander are lost:  Nest site fidelity in the large carpenter bee, Xylocopa virginica. Insectes Sociaux 58, 127-133.

Rehan SM, MH Richards. 2010. The influence of maternal quality on brood sex allocation in the small carpenter bee, Ceratina calcarata.  Ethology 116, 876-887.

Rehan SM, TW Chapman, AI Craigie, MH Richards, SJB Cooper, MP Schwarz. 2010. Molecular phylogeny of the small carpenter bees (Hymenoptera: Apidae: Ceratinini) indicates early and rapid global dispersal.  Molecular Phylogenetics and Evolution 55, 1042-1054

Skandalis A, M Frampton, J Seger, MH Richards. 2010. Alternative splicing of polB in primates. RNA 16, 2014–2022.

Richards MH, J Vickruck, SM Rehan. 2010. Colony social organisation of Halictus confusus in southern Ontario, with comments on sociality in the subgenus H. (Seladonia). Journal of Hymenoptera Research 19, 144-158.


Rehan SM, MH Richards, MP Schwarz. 2009. Evidence of social nesting in the Ceratina of Borneo. Journal of the Kansas Entomological Society 82, 194-209.

Richardson JML, MH Richards. 2008. A randomization program to compare species richness values.  Insect Conservation and Diversity 1, 135-141.

Rehan SM, MH Richards. 2008. Morphological and DNA sequence delineation of two problematic species of Ceratina (Hymenoptera: Apidae) from Eastern Canada. Journal of the Entomological Society of Ontario 139, 59-67.

Skandalis DA, GJ Tattersall, S Prager, and MH Richards. 2008. Local and latitudinal contributions to size variation of the large carpenter bee, Xylocopa virginica. Journal of the Kansas Entomological Society 82, 30-42.

Frampton M, S Droege, T Conrad, S Prager, MH Richards. 2008. Evaluation of specimen preservatives for DNA analyses of bees. Journal of Hymenoptera Research 17, 195-200.

Long ago…

Schwarz MP, MH Richards, BN Danforth. 2007. Changing paradigms in insect social evolution: insights from halictine and allodapine bees. Annual Review of Entomology 52, 127-150.

Gray DA, P Barnfield, M Seifried, MH Richards. 2006. Molecular divergence between Gryllus rubens and Gryllus texensis, sister species of field crickets (Orthoptera: Gryllidae). The Canadian Entomologist 138, 305-313.  PDF

Richards MH. 2004. Annual and social variation in foraging effort of the obligately eusocial sweat bee, Halictus ligatus Say (Hymenoptera, Halictidae). Journal of the Kansas Entomological Society, 77 (4), 484-502. PDF

Wyman LM, MH Richards. 2003.  Colony social organization of Lasioglossum malachurum Kirby (Hymenoptera; Halictidae) in southern Greece. Insectes Sociaux 50, 201-211.  PDF

Richards MH. 2000. Evidence for geographic variation in colony social organization in an obligately social sweat bee, Lasioglossum malachurum Kirby (Hymnoptera; Halictidae).  Canadian Journal of Zoology 78, 1259-1266.  PDF

Richards MH. 1994.  Social evolution in the genus Halictus: A phylogenetic approach. Insectes Sociaux 41, 315-325.  PDF


Leave a Reply

Fill in your details below or click an icon to log in: Logo

You are commenting using your account. Log Out /  Change )

Google+ photo

You are commenting using your Google+ account. Log Out /  Change )

Twitter picture

You are commenting using your Twitter account. Log Out /  Change )

Facebook photo

You are commenting using your Facebook account. Log Out /  Change )


Connecting to %s